xtrabackup备份MySQL报错：InnoDB: Error number 24 means 'Too many open files'

190228 00:30:01 >> log scanned up to (88343615848)
xtrabackup: Generating a list of tablespaces
InnoDB: Allocated tablespace ID 2 for mysql/plugin, old maximum was 0
InnoDB: Operating system error number 24 in a file operation.
InnoDB: Error number 24 means ‘Too many open files‘
InnoDB: Some operating system error numbers are described at http://dev.mysql.com/doc/refman/5.7/en/operating-system-error-codes.html
InnoDB: File ./jira/AO_4B00E6_STASH_SETTINGS.ibd: ‘open‘ returned OS error 124. Cannot continue operation
InnoDB: Cannot continue operation.

[root@linux mysql]# ulimit -a
core file size          (blocks, -c) 0
data seg size           (kbytes, -d) unlimited
scheduling priority             (-e) 0
file size               (blocks, -f) unlimited
pending signals                 (-i) 385500
max locked memory       (kbytes, -l) 64
max memory size         (kbytes, -m) unlimited
open files                      (-n) 1024
pipe size            (512 bytes, -p) 8
POSIX message queues     (bytes, -q) 819200
real-time priority              (-r) 0
stack size              (kbytes, -s) 8192
cpu time               (seconds, -t) unlimited
max user processes              (-u) 385500
virtual memory          (kbytes, -v) unlimited
file locks                      (-x) unlimited

vim /etc/security/limits.conf

* soft nofile 65535

* hard nofile 65535

vim /etc/my.cnf

innodb_open_files=1000